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1. Leader cells in epithelial repair are directed by p53, which activates the cell cycle inhibitor p21WAF1/CIP1 to induce leader behavior through inhibition of cyclin-dependent kinase activity.

2. P53 also induces crowding hypersensitivity in leader cells, ensuring their clearance upon epithelial repair through cell competition.

3. Mechanically induced p53 directs the emergence of a transient population of leader cells that drive migration and ensure their clearance upon epithelial repair.

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